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Data availability All data generated or analyzed during this study are included in this published article

The dual mechanism produces additive or combined effects on energy balance -- GIPR activation in the CNS may enhance the anorectic signals from GLP-1R activation, and peripheral GIPR signaling improves adipose tissue function and lipid handling in ways GLP-1 alone does not

FOXO4-DRI treatment improved the testicular microenvironment and partially restored testosterone levels in naturally aged mice

Central Nervous System Pathways: Cag acts centrally through the brain rather than purely on peripheral tissues