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Under a lower red/far-red ratio, we propose that an increase in ROS and H 2 O 2 in chloroplasts arises from inefficient chlorophyll biosynthesis in seedlings after transition to white light or in mature shaded leaves from an imbalance in the excitation state of the two photosystems (short term) and limited availability of reductants and assimilates (long term), which again initiates the accumulation and redistribution of glutathione (and ascorbate) in plant cell compartments (Fig

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